Tuesday, March 3, 2009

Jellyfish

Jellyfish (also known as jellies or sea jellies) are free-swimming members of the phylum Cnidaria. They have several different morphologies that represent several different cnidarian classes including the Scyphozoa (about 200 species), Staurozoa (about 50 species), Cubozoa (about 20 species), and Hydrozoa (about 1000-1500 species that make jellyfish and many more that do not). The jellyfish in these groups are also called, respectively, scyphomedusae, stauromedusae, cubomedusae, and hydromedusae; "medusa" (plural "medusae") is another word for jellyfish. Jellyfish are found in every ocean, from the surface to the deep sea. Some hydrozoan jellyfish, or hydromedusae, are also found in fresh water. Most of the information about jellyfish that follows in this article is about scyphozoan jellyfish, or scyphomedusae. These are the big, often colorful, jellyfish that are common in coastal zones worldwide.

In its broadest sense, the term jellyfish is sometimes used also to refer to members of the phylum Ctenophora. Although not closely related to cnidarian jellyfish, ctenophores are also free-swimming planktonic carnivores, are also generally transparent or translucent, and occur in shallow to deep portions of all the world's oceans. Ctenophores move using eight rows of fused cilia that beat in metachronal waves that diffract light, so that they sparkle with all of the colors of the rainbow. The rest of this article deals only with jellyfish in the phylum Cnidaria.

Jellyfish don't have specialized digestive, osmoregulatory, central nervous, respiratory, or circulatory systems. They digest using the gastrodermal lining of the gastrovascular cavity, where nutrients are absorbed. They do not need a respiratory system since their skin is thin enough that the body is oxygenated by diffusion. They have limited control over movement, but can use their hydrostatic skeleton to accomplish movement through contraction-pulsations of the bell-like body; some species actively swim most of the time, while others are passive much of the time. Jellyfish are composed of more than 90% water; most of their umbrella mass is a gelatinous material - the jelly - called mesoglea which is surrounded by two layers of epithelial cells which form the exumbrella (top surface) and subumbrella (bottom surface) of the bell, or body.

Jellyfish do not have a brain or central nervous system, but rather have a loose network of nerves, located in the epidermis, which is called a "nerve net." A jellyfish detects various stimuli including the touch of other animals via this nerve net, which then transmits impulses both throughout the nerve net and around a circular nerve ring, through the rhopalial lappet, located at the rim of the jellyfish body, to other nerve cells. Some jellyfish also have ocelli: light-sensitive organs that do not form images but which can detect light, and are used to determine up from down, responding to sunlight shining on the water's surface.

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